Fisiologia Vegetal.indb

Propaganda
Aula
NUTRIÇÃO MINERAL
DAS PLANTAS
4
META
(VWXGDUDVIXQo}HVGRVPLQHUDLVQDQXWULomRGDVSODQWDV
OBJETIVOS
$R¿QDOGHVWDDXODRDOXQRGHYHUi
FRPSUHHQGHURVHOHPHQWRVHVVHQFLDLVPDFURHPLFURQXWULHQWHVHVXDVIXQo}HVQDSODQWD
35e5(48,6,726
Conceitos de absorção de água e sais minerais na planta.
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Fisiologia Vegetal
INTRODUÇÃO
$VSODQWDVVmRRUJDQLVPRVDXWRWUyÀFRVTXHYLYHPHQWUHGRLVDPELHQWHV
LQWHLUDPHQWHLQRUJkQLFRVUHWLUDQGR&2GDDWPRVIHUDHiJXDHQXWULHQWHV
PLQHUDLVGRVROR2VQXWULHQWHVPLQHUDLVVmRDGTXLULGRVSULPDULDPHQWHQD
IRUPDGHtRQVLQRUJkQLFRVHHQWUDPQDELRVIHUDSUHGRPLQDQWHPHQWHDWUDYpV
GRVLVWHPDUDGLFXODUGDSODQWD$JUDQGHiUHDVXSHUÀFLDOGDVUDt]HVHVXD
JUDQGHFDSDFLGDGHSDUDDEVRUYHUtRQVLQRUJkQLFRVHPEDL[DVFRQFHQWUDo}HV
QDVROXomRGRVRORWRUQDPDDEVRUomRPLQHUDOSHODSODQWDXPSURFHVVR
bastante efetivo. Além disso, outros organismos, como os fungos (micorUt]LFRVHDVEDFWpULDVÀ[DGRUDVGHQLWURJrQLRIUHTXHQWHPHQWHFRQWULEXHP
SDUDDDTXLVLomRGHQXWULHQWHVSHODVSODQWDV'HSRLVGHDEVRUYLGRVRVtRQV
VmRWUDQVSRUWDGRVSDUDDVGLYHUVDVSDUWHVGDSODQWDRQGHVmRDVVLPLODGRV
HXWLOL]DGRVHPLPSRUWDQWHVIXQo}HVELROyJLFDV
2 HVWXGR GH FRPR DV SODQWDV DEVRUYHP WUDQVSRUWDP DVVLPLODP
H XWLOL]DP RV tRQV p FRQKHFLGR FRPR 1875,d®2 0,1(5$/ (VWD
área do conhecimento busca o entendimento das relações iônicas sob
condições naturais de solo (salinidade, acidez, alcalinidade, presença de
HOHPHQWRVWy[LFRVFRPRDOXPtQLRHPHWDLVSHVDGRVHWFSRUpPRVHX
maior interesse está ligado diretamente à agricultura e à produtividade
GDVFXOWXUDV$OWDSURGXomRDJUtFRODGHSHQGHIRUWHPHQWHGDIHUWLOL]DomR FRP HOHPHQWRV PLQHUDLV 1R HQWDQWR DV SODQWDV FXOWLYDGDV
tipicamente, utilizam menos da metade dos fertilizantes aplicados.
2UHVWDQWHSRGHVHUOL[LYLDGRSDUDRVOHQoyLVVXEWHUUkQHRVGHiJXD
WRUQDUVHÀ[DGRDRVRORRXFRQWULEXLUSDUDDSROXLomRGRDU$VVLP
WRUQDVHGHJUDQGHLPSRUWkQFLDDXPHQWDUDHÀFLrQFLDGHDEVRUomRH
GHXWLOL]DomRGHQXWULHQWHVUHGX]LQGRRVFXVWRVGHSURGXomRHFRQWULEXLQGRSDUDHYLWDUSUHMXt]RVDRPHLRDPELHQWH
)HUWLOL]DomR GD WHUUD FRP ILQV DJUtFRODV XWLOL]DQGR FRPSRVWRV
orgânicos (Fonte: http://www.guiasjp.com).
34
Nutrição mineral das plantas
SOLO
2FRQFHLWRGHVRORFRPRPHLRSDUDRFUHVFLPHQWRYHJHWDOpXPDQRomR
TXHYHPGHVGHRVSULPyUGLRVGDDJULFXOWXUD'HIDWRDVFDUDFWHUtVWLFDVItVLFDV
HTXtPLFDVGRVVRORVFRQGLFLRQDPRFUHVFLPHQWRYHJHWDODRID]HUYDULDU
DFDSDFLGDGHGHUHWHQomRGHiJXDDVROXELOLGDGHGRVHOHPHQWRVPLQHUDLV
DVWUDQVIRUPDo}HVPLQHUDLVHELRTXtPLFDVDOL[LYLDomRGRVQXWULHQWHVHR
S+2VRORpLPSRUWDQWHSDUDRFUHVFLPHQWRYHJHWDOSRLVVXSUHDVSODQWDV
FRP IDWRUHV GH FUHVFLPHQWR SHUPLWH R GHVHQYROYLPHQWR H GLVWULEXLomR
GDVVXDVUDt]HVHSRVVLELOLWDRPRYLPHQWRGRVQXWULHQWHVGHiJXDHDUQDV
VXSHUItFLHVUDGLFXODUHV
2VRORpXPVLVWHPDFRPSOH[RFRQVWLWXtGRGHIDVHVPDWUL]GRVROR
VROXomRGRVRORHDIDVHJDVRVD
$IDVHVyOLGDGRVRORPDWUL]pFRQVWLWXtGDSHODVIUDo}HVPLQHUDOHRUJkQLFD
$IUDomRPLQHUDOUHVXOWDGDDomRGDLQWHPSHUL]DomRItVLFDTXtPLFDHELROyJLFD
VREUHDVURFKDVHpFRQVWLWXtGDGHSDUWtFXODVGHGLIHUHQWHVWDPDQKRV
DUHLDJURVVDPPGHGLkPHWUR
DUHLDÀQDPPGHGLkPHWUR
VLOWHPPGHGLkPHWUR
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)UDFLRQDPHQWR SURSRVWR SHOD &RPLVVmR 3HUPDQHQWH GH 0pWRGRV GH
&DPSRGD6%&6)
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RVHOHPHQWRVHVVHQFLDLVeFRQVWLWXtGDGHXPDVROXomRGHHOHWUyOLWRVHP
HTXLOtEULRFRPDIDVHVyOLGD
2DUGRVRORWHPJHUDOPHQWHRVPHVPRVFRPSRQHQWHVGRDUDWPRVIpULFR$UHVSLUDomRGDVUDt]HVHGRVPLFURUJDQLVPRVDGHFRPSRVLomR
GDPDWpULDRUJkQLFDHRXWUDVUHDo}HVSRUpPPRGLÀFDPVXDFRPSRVLomR
Aula
4
$XWRWUy¿FRV
É o nome dado à
qualidade do ser
vivo de produzir
seu próprio alimento.
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ELEMENTOS ESSENCIAIS
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quantidades, variáveis conforme a espécie e o estado de desenvolvimento.
(VVHVHOHPHQWRVHQFRQWUDPVHQRVVRORVHPGLIHUHQWHVFRPELQDo}HVTXtPLFDVVHQGRVyDOJXPDVGHVWDVSDVVtYHLVGHVHUHPDEVRUYLGDVSHODVSODQWDV
2SURJUHVVRQDTXtPLFDDQDOtWLFDHVSHFLDOPHQWHRGHVHQYROYLPHQWRGH
35
Fisiologia Vegetal
WpFQLFDVGHSXULÀFDomRGHVDLVHGHWHUPLQDomRGHHOHPHQWRVPLQHUDLVHP
quantidades traços, associado ao desenvolvimento de técnicas de cultivo
GHSODQWDHPVROXomRQXWULWLYDSHUPLWLXRHVWDEHOHFLPHQWRGRVFULWpULRV
de essencialidade.
Para um elemento ser considerado essencial, deve satisfazer a três
critérios básicos:
D$SODQWDQmRSRGHVHUFDSD]GHFRPSOHWDUVHXFLFOR´YLWDOµQDDXVrQFLD
do elemento mineral.
E$IXQomRGHFHUWRHOHPHQWRPLQHUDOQmRSRGHVHUVXEVWLWXtGRSRURXWUR
elemento mineral.
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SODQWDRXVHUUHTXHULGRQXPDGHWHUPLQDGDHWDSDPHWDEyOLFD
$VSODQWDVVXSHULRUHVUHTXHUHPDOpPGR&+H2WUH]HHOHPHQWRVTXH
HODVDEVRUYHPQDIRUPDGHtRQVGDVROXomRGRVROR6HLVGHVWHVUHTXHULGRV
HP PDLRUHV TXDQWLGDGHV VmR FKDPDGRV 0$&521875,(17(6 1 3
.&D6H0J2VVHWHRXWURVUHTXHULGRVHPEDL[DVFRQFHQWUDo}HVVmR
FKDPDGRV0,&521875,(17(6)H0Q&X=Q%0RH&O7DEHOD
7DEHOD(OHPHQWRVPLQHUDLVHVVHQFLDLVHEHQpÀFRV
para plantas superiores e inferiores.
2VGHQRPLQDGRVHOHPHQWRV´EHQpÀFRVµVmRDTXHOHVPLQHUDLVTXH
&RPSHQVDPRXHOLPLQDPRVHIHLWRVWy[LFRVGHRXWURV([2$OHP
FRQFHQWUDomRDEDL[RGHSSPSRGHUHGX]LURXHOLPLQDUHIHLWRVWy[LFRV
GH&X0QH3
6XEVWLWXHPXPHOHPHQWRHVVHQFLDOHPDOJXPDGHVXDVIXQo}HVPHQRV
HVSHFtÀFDV([21DSRGHVDWLVID]HUSDUWHGDIXQomRRVPyWLFDGR.
6mRHVVHQFLDLVDSHQDVSDUDDOJXPDVHVSpFLHV([21DpHVVHQFLDOSDUD
DKDOyÀWDAtriplex vesicaria.
2VHOHPHQWRVPLQHUDLVPDFURHPLFURQXWULHQWHVDRODGRGHIDWRUHVWDLV
como luz, água e gás carbônico constituem a matéria prima que a maquinaria biossintética da célula utiliza para crescer e se desenvolver. Embora
36
Nutrição mineral das plantas
FRQVWLWXDPDSHQDVGHDGDPDWpULDVHFDWRWDORVHOHPHQWRVPLQHUDLV
além de serem componentes das moléculas essenciais, constituem estruturas
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RVPyWLFRWUDQVSRUWHGHHOpWURQVVLVWHPDWDPSmRGRSURWRSODVPDHFRQWUROH
de permeabilidade etc.
2V QXWULHQWHV VmR QHFHVViULRV SDUD D SODQWD OHYDU D FDER GHWHUPLQDGDV
IXQo}HV SHOR TXH D FDUrQFLD H R H[FHVVRHVWmRUHODFLRQDGRVFRPVLQWRPDV
YLVtYHLVTXHSRUVXDYH]HVWmRUHODFLRQDGRVFRPDVXDPRELOLGDGHHIXQomR
2VVLQWRPDVPDLVFRPXQVRFRUUHPDRQtYHOGDIRUPDomRGRVQyVGDLQVHUomR
foliar, do crescimento vegetal, do desenvolvimento do sistema radicular (cor,
grau de desenvolvimento) e das folhas que, se apresentarem manchas amarelas,
têm clorose e, se apresentarem manchas negras, têm necrose.
Aula
4
FUNÇÕES DOS MINERAIS NAS PLANTAS
6mRPDFURQXWULHQWHV136&D0JH.
21LWURJrQLR1HQWUDSULQFLSDOPHQWHQDFRQVWLWXLomRGHFRPSRVWRV
RUJkQLFRVVHQGRXPQXWULHQWHPyYHO(PH[FHVVRSURYRFDXPFUHVFLmento vegetal acelerado, originando folhas de cor verde-escura, ocorre
XPDGLPLQXLomRGDUHVLVWrQFLDDGRHQoDVXPUHWDUGDPHQWRGDÁRUDomRH
o ciclo de vida é encurtado. A carência de nitrogênio reduz o crescimento
IROLDUSURYRFDDFORURVHIROLDU2VUDPRVFDXOLQDUHVÀFDPS~USXUDVRX
vermelhos, localizando-se inicialmente os sintomas em partes velhas da
planta.
2)yVIRUR3WDPEpPLQWHUYpPQDIRUPDomRGHFRPSRVWRVRUJkQLFRV
HVSHFLDOPHQWH$73HIRVIROtSLGRVVHQGRXPQXWULHQWHPyYHO$FDUrQFLDGH
IyVIRURUHGX]RFUHVFLPHQWRFDXOLQDUHUDGLFXODUHSURYRFDRDSDUHFLPHQWR
GHiUHDVQHFUyWLFDVQDVIROKDVHSHFtRORVDVFpOXODVGHL[DUmRGHID]HURVHX
PHWDEROLVPRHPRUUHUmR$VIROKDVMRYHQVWrPWHQGrQFLDSDUDHVFXUHFHURX
ÀFDUYHUGHD]XODGDVHQTXDQWRTXHDVPDLVYHOKDVÀFDPYHUPHOKDV1XPD
fase inicial, os sintomas acentuam-se nas partes mais velhas da planta.
2(Q[RIUH6LQWHUYpPQDVtQWHVHGHFRPSRVWRVRUJkQLFRVHPHVSHFLDO
YLWDPLQDVHHQ]LPDVVHQGRXPQXWULHQWHLPyYHO1mRVHFRQKHFHVLQWRPDWRORJLDSDUDRVHXH[FHVVR$FDUrQFLDGHHQ[RIUHUHGX]RFUHVFLPHQWR
YHJHWDOSURYRFDQGRDFORURVHIROLDU,QLFLDOPHQWHRVVLQWRPDVDFHQWXDPVH
nas zonas mais jovens da planta.
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PHULVWHPiWLFDVVHQGRXPQXWULHQWHLPyYHO(PH[FHVVRDOWHUDRULWPRGD
GLYLVmRFHOXODU$FDUrQFLDGHFiOFLRRULJLQDPDOIRUPDo}HVQDVIROKDVMRYHQV
FXUYDPHQWRGRViSLFHVFORURVHPDUJLQDOTXHSURJULGHSDUDQHFURVHUHGXomR
GRFUHVFLPHQWRUDGLFXODUHPXGDQoDGDFRORUDomRGDVUDt]HVSDUDFDVWDQKR
37
Fisiologia Vegetal
,QLFLDOPHQWHRVVLQWRPDVDFHQWXDPVHQDV]RQDVPDLVMRYHQVGDVSODQWDV
20DJQpVLR0JpXPFRQVWLWXLQWHGDFORURÀODHGDVSURWHtQDVEHPFRPR
de cofatores enzimáticos, sendo essencial ao funcionamento dos ribossomas.
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de cálcio e potássio. A carência de magnésio provoca cloroses intervenais,
QHFURVHIROLDUHQFXUWDPHQWRGHHQWUHQyVUHGXomRGRFUHVFLPHQWRYHJHWDO
LQLELomRGDÁRUDomRPRUWHSUHPDWXUDGDVIROKDVHGHJHQHUDomRGRVIUXWRV
,QLFLDOPHQWHRVVLQWRPDVDFHQWXDPVHQDV]RQDVPDLVYHOKDVGDVSODQWDV
23RWiVVLR.pXPUHJXODGRURVPyWLFRQHFHVViULRjDWLYLGDGHHQ]LPiWLFDHjVtQWHVHSURWHLFDVHQGRXPQXWULHQWHPyYHO1mRVHFRQKHFHVLQWRPDWRORJLDSDUDRVHXH[FHVVR$FDUrQFLDGHSRWiVVLRSURYRFDXPFUHVFLPHQWR
YHJHWDOPXLWRUHGX]LGRFORURVHPDWL]DGDGDIROKDPDQFKDVQHFUyWLFDVIROKDV
UHFXUYDGDVHHQURODGDVVREUHDIDFHVXSHULRUHHQFXUWDPHQWRGHHQWUHQyV
,QLFLDOPHQWHRVVLQWRPDVDFHQWXDPVHQDV]RQDVPDLVYHOKDVGDVSODQWDV
2EV2&iOFLR&D(Q[RIUH6HR0DJQpVLR0JHPPXLWRVFDVRV
VmRHQFRQWUDGRVHGHQRPLQDGRVFRPRPDFURQXWULHQWHVVHFXQGiULRV
6mRPLFURQXWULHQWHV)H&X0Q=Q0RH%H&O
2 )HUUR )H p XP FRQVWLWXLQWH GR JUXSR SURVWpWLFR GH SURWHtQDV
QHFHVViULR j VtQWHVH GH FORURÀOD H j GLYLVmR FHOXODU VHQGR XP QXWULHQWH
LPyYHO1mRVHFRQKHFHVLQWRPDWRORJLDSDUDRVHXH[FHVVR$FDUrQFLDGH
IHUURSURYRFDXPDH[WHQVDFORURVHIROLDU
2 &REUH &X p XP DFHLWDGRU LQWHUPHGLiULR GH HOpWURQV VHQGR XP
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carência de cobre altera a tonalidade das folhas, tornando-as verde-azuladas
e enroladas onde aparecem cloroses intervenais e necroses.
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R[LUUHGXomR HVVHQFLDLV j IRWRVVtQWHVH H j VtQWHVH GH FORURÀOD VHQGR XP
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carência de manganês provoca clorose intervenal nas zonas mais jovens,
enrolamento e queda de folhas.
2=LQFR=QpXPDDWLYDGRUHQ]LPiWLFRVHQGRXPQXWULHQWHPyYHO
1mRVHFRQKHFHVLQWRPDWRORJLDSDUDRVHXH[FHVVR$FDUrQFLDGH]LQFR
SURYRFDXPDUHGXomRGRFUHVFLPHQWRYHJHWDO
2 0ROLEGpQLR 0R p HVVHQFLDO SDUD D À[DomR GH QLWURJrQLR H
DVVLPLODomRGHQLWUDWRVVHQGRXPQXWULHQWHLPyYHO1mRVHFRQKHFHVLQWRPDWRORJLDSDUDRVHXH[FHVVR$FDUrQFLDGHPROLEGpQLRRULJLQDPDQFKDV
FORUyWLFDVLQWHUYHQDLVVHJXLGDVGHQHFURVHPDUJLQDOHHQURODPHQWRIROLDU
LQWHUIHULQGRFRPDIUXWLÀFDomR
2%RUR%pXPUHJXODGRUGHPHWDEROLVPRQHFHVViULRjWUDQVORFDomR
GHDo~FDUHVVHQGRXPQXWULHQWHLPyYHO1mRVHFRQKHFHVLQWRPDWRORJLD
SDUDRVHXH[FHVVR$FDUrQFLDGHERURDIHWDRVyUJmRVGHUHVHUYD
2&ORUR&OpQHFHVViULRjIRWRVVtQWHVHVHQGRXPQXWULHQWHPyYHO(P
H[FHVVRSURYRFDFORURVHHQHFURVHIROLDU$FDUrQFLDGHFORURUHGX]RFUHVFLmento vegetal em mais de 50 % e provoca o aparecimento de folhas murchas.
38
Nutrição mineral das plantas
CONCLUSÃO
Aula
4
A terra é formada de 90 elementos de ocorrência natural, sendo os mais
FRPXQVRR[LJrQLRVLOtFLRDOXPtQLRHIHUUR2VHOHPHQWRVVmRHQFRQWUDGRV
QRVRORVREDIRUPDGHPLQHUDLVHXWLOL]DGRVSHODVSODQWDVSDUDUHDOL]DomRGR
seu metabolismo. Sabe-se que as plantas superiores requerem um total de
16 elementos para seu crescimento normal. Destes, o carbono, o hidrogênio
HRR[LJrQLRSURYrPGRDUHGDiJXD2UHVWDQWHpDEVRUYLGRSHODVUDt]HVVRE
DIRUPDGHtRQV(VWHVHOHPHQWRVVmRFODVVLÀFDGRVHPPDFURQXWULHQWHVH
PLFURQXWULHQWHV2VHOHPHQWRVPLQHUDLVGHVHPSHQKDPQXPHURVDVIXQo}HV
importantes nas células. Regulam a osmose e afetam a permeabilidade celular.
Alguns servem como receptores de elétrons, como componentes estruturais
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ponentes estruturais de enzima.
RESUMO
Nesta aula você pode observar que as plantas utilizam diversos minerais
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HjVHLYDHODERUDGDQDIRUPDomRGRVFRPSRQHQWHVFHOXODUHV
2VQXWULHQWHVPLQHUDLVVmRDGTXLULGRVSULPDULDPHQWHQDIRUPDGHtRQV
inorgânicos e entram na biosfera, predominantemente, através do sistema
UDGLFXODUGDSODQWD$JUDQGHiUHDVXSHUÀFLDOGDVUDt]HVHVXDJUDQGHFDSDFL
GDGHSDUDDEVRUYHUtRQVLQRUJkQLFRVHPEDL[DVFRQFHQWUDo}HVQDVROXomRGR
VRORWRUQDPDDEVRUomRPLQHUDOSHODSODQWDXPSURFHVVREDVWDQWHHIHWLYR
$VFDUDFWHUtVWLFDVItVLFDVHTXtPLFDVGRVVRORVFRQGLFLRQDPRFUHVFLPHQ
WRYHJHWDODRID]HUYDULDUDFDSDFLGDGHGHUHWHQomRGHiJXDDVROXELOLGDGHGRV
HOHPHQWRVPLQHUDLVDVWUDQVIRUPDo}HVPLQHUDLVHELRTXtPLFDVDOL[LYLDomR
GRVQXWULHQWHVHRS+2VRORpLPSRUWDQWHSDUDRFUHVFLPHQWRYHJHWDOSRLV
supre as plantas com fatores de crescimento, permite o desenvolvimento
HGLVWULEXLomRGDVVXDVUDt]HVHSRVVLELOLWDRPRYLPHQWRGRVQXWULHQWHVGH
iJXDHDUQDVVXSHUItFLHVUDGLFXODUHV
2VQXWULHQWHVHVVHQFLDLVVmRUHTXHULGRVSHODVSODQWDVHPGHWHUPL
nadas quantidades, variáveis conforme a espécie e o estado de desenvolvimento. Esses elementos encontram-se nos solos em diferentes combiQDo}HVTXtPLFDVVHQGRVyDOJXPDVGHVVDVSDVVtYHLVGHVHUHPDEVRUYLGDV
pelas plantas.
2V QXWULHQWHV TXH VmR H[LJLGRV HP PDLRUHV TXDQWLGDGHV VmR GHQR
PLQDGRVPDFURQXWULHQWHVHRVTXHVmRH[LJLGRVHPEDL[DVFRQFHWUDo}HV
denomina-se micronutrientes.
39
Fisiologia Vegetal
ATIVIDADES
1. Descreva a origem dos elementos minerais.
COMENTÁRIO SOBRE A ATIVIDADE
2 VROR UHSUHVHQWD R SURGXWR ÀQDO GH IHQ{PHQRV ItVLFRV TXtPLFRV H
ELROyJLFRVHFRQVWLWXLDSRUomRGDFURVWDWHUUHVWUHRQGHDVSODQWDVFUHVFHP
PRÓXIMA AULA
1DSUy[LPDDXODLUHPRVDERUGDUDQXWULomRPLQHUDOGDVSODQWDV
AUTOAVALIAÇÃO
5HVSRQGDDVTXHVW}HVDEDL[R
2TXHVmRHOHPHQWRVHVVHQFLDLV"
4XDODIXQomRGRSRWiVVLRQDVSODQWDVVXSHULRUHV"
4XDORVHOHPHQWRVPLQHUDOLVTXHQmRpREWLGRDSDUWLUGRVROR"
REFERÊNCIAS
)(55(,5$/*5Fisiologia vegetal: UHODo}HVKtGULFDVHG)RUWDOH]D
(GLo}HV8)&
+23.,16:*Introduction to plant physiologyHG1HZ<RUN
-RKQ:LOH\6RQV,QF
3859(6:.HWDOVida: a ciência da biologia. 6 ed. Porto Alegre: ArtPHG
5$9(13+(9(575)(,&++2516(Biologia vegetal. 6 ed.
5LRGH-DQHLUR*XDQDEDUD.RRJDQ
7$,=/=(,*(5(Fisiologia VegetalHG(GLWRUD$UWPHG
40
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